Marine biology

Marine biology is the scientific study of living organisms in the ocean or other marine or brackish bodies of water. Given that in biology many phyla, families and genera have some species that live in the sea and others that live on land, marine biology classifies species based on the environment rather than on taxonomy.

Marine Life is a vast resource, providing food, medicine, and raw materials, in addition to helping to support recreation and tourism all over the world. At a fundamental level, marine life helps determine the very nature of our planet. Marine organisms contribute significantly to the oxygen cycle, and are involved in the regulation of the earth's climate.Shorelines are in part shaped and protected by marine life, and some marine organisms even help create new land.

Marine biology covers a great deal, from the microscopic, including plankton and phytoplankton, which can be as small as 0.02 micrometers and are both hugely important as the primary producers of the sea, to the huge cetaceans (whales) which reach up to a reported 48 meters (125 feet) in length.

The habitats studied by marine biology include everything from the tiny layers of surface water in which organisms and abiotic items may be trapped in surface tension between the ocean and atmosphere, to the depths of the abyssal trenches, sometimes 10,000 meters or more beneath the surface of the ocean. It studies habitats such as coral reefs, kelp forests, tidepools, muddy, sandy, and rocky bottoms, and the open ocean (pelagic) zone, where solid objects are rare and the surface of the water is the only visible boundary.

A large amount of all life on Earth exists in the oceans. Exactly how large the proportion is still unknown. While the oceans comprise about 71% of the Earth's surface, due to their depth they encompass about 300 times the habitable volume of the terrestrial habitats on Earth.

Many species are economically important to humans, including the food fishes. It is also becoming understood that the well-being of marine organisms and other organisms are linked in very fundamental ways. Human understanding is growing of the relationship between life in the sea and important cycles such as that of matter (such as the carbon cycle) and of air (such as Earth's respiration, and movement of energy through ecosystems). Large areas beneath the ocean surface still remain effectively unexplored.

Albatross

Albatrosses, of the biological family Diomedeidae, are large seabirds allied to the procellariids, storm-petrels and diving-petrels in the order Procellariiformes (the tubenoses). They range widely in the Southern Ocean and the North Pacific. They are absent from the North Atlantic, although fossil remains show they once occurred there too and occasional vagrants turn up. Albatrosses are amongst the largest of flying birds, and the great albatrosses (genus Diomedea) have the largest wingspans of any extant birds. The albatrosses are usually regarded as falling into four genera, but there is disagreement over the number of species.

Albatrosses are highly efficient in the air, using dynamic soaring and slope soaring to cover great distances with little exertion. They feed on squid, fish and krill by either scavenging, surface seizing or diving. Albatrosses are colonial, nesting for the most part on remote oceanic islands, often with several species nesting together. Pair bonds between males and females form over several years, with the use of 'ritualised dances', and will last for the life of the pair. A breeding season can take over a year from laying to fledging, with a single egg laid in each breeding attempt.

Of the 21 species of albatrosses recognised by the IUCN, 19 are threatened with extinction. Numbers of albatrosses have declined in the past due to harvesting for feathers, but today the albatrosses are threatened by introduced species such as rats and feral cats that attack eggs, chicks and nesting adults; by pollution; by a serious decline in fish stocks in many regions largely due to overfishing; and by long-line fishing. Long-line fisheries pose the greatest threat, as feeding birds are attracted to the bait, become hooked on the lines, and drown. Identified stakeholders such as governments, conservation organisations and people in the fishing industry are all working towards reducing this bycatch.

Annelid

The annelids, collectively called Annelida (from Latin anellus "little ring"), are a large phylum of animals comprising the segmented worms, with about 15,000 modern species including the well-known earthworms and leeches. They are found in most wet environments, and include many terrestrial, freshwater, and especially marine species (such as the polychaetes), as well as some which are parasitic or mutualistic. They range in length from under a millimeter to over 3 meters (the seep tube worm Lamellibrachia luymesi).

Annelids are bilaterally symmetric and triploblastic protostomes with a coelom (which makes them coelomates), closed circulatory system and true segmentation. Their segmented bodies and coelom have given them evolutionary advantages over other worms. Oligochaetes and polychaetes typically have spacious coeloms; in leeches, the coelom is filled in with tissue and reduced to a system of narrow canals; archiannelids may lack the coelom entirely. The coelom is divided into a sequence of compartments by walls called septa.

In the most general forms each compartment corresponds to a triple segment of the body, which also includes a portion of the nervous and (closed) circulatory systems, allowing it to function relatively independently. The closed circulatory system consists of networks of vessels containing blood with oxygen-carrying hemoglobin. Dorsal and ventral vessels are connected by segmental pairs of vessels. The dorsal vessel and five pairs of vessels that circle the esophagus of an earthworm are muscular and pump blood through the circulatory system.

Tiny blood vessels are abundant in the earthworm's skin, which function as its respiratory organ. Each segment (metamere) is marked externally by one or more rings, called annuli. Each segment also has an outer layer of circular muscle underneath a thin cuticle and epidermis, and a system of longitudinal muscles. In earthworms and in daria the longitudinal muscles are strengthened by collagenous lamellae; the leeches have a double layer of muscles between the outer circulars and inner longitudinals. In most forms they also carry a varying number of bristles, called setae, and among the polychaetes a pair of appendages, called parapodia.

Anterior to the true segments lies the prostomium and peristomium, which carries the mouth, and posterior to them lies the pygidium, where the anus is located. The digestive tract is quite variable but is usually specialized. For example, in some groups (notably most earthworms) it has a typhlosole (to increase surface area) along much of its length. Different species of annelids have a wide variety of diets, including active and passive hunters, scavengers, filter feeders, direct deposit feeders which simply ingest the sediments, and blood-suckers.

The vascular system and the nervous system are separate from the digestive tract. The vascular system includes a dorsal vessel conveying the blood toward the front of the worm, and a ventral longitudinal vessel which conveys the blood in the opposite direction. The two systems are connected by a vascular sinus and by lateral vessels of various kinds, including in the true earthworms, capillaries on the body wall.

The nervous system has a nerve cord from which lateral nerves come in contact with each segment. Every segment has an autonomy; however, they unite to perform as a single body for functions such as locomotion. Growth in many groups occurs by replication of individual segmental units, in others the number of segments is fixed in early development.

Tunicate

Tunicate, also known as urochordata, tunicata (and by the common names of urochordates, sea squirts, and sea pork is the subphylum of a group of underwater saclike filter feeders with incurrent and excurrent siphons, that are members of the phylum Chordata. Most tunicates feed by filtering sea water through pharyngeal slits, but some are sub-marine predators such as the Megalodicopia hians.

Like other chordates, tunicates have a notochord during their early development, but lack segmentation throughout the body and tail. Tunicates lack the kidney-like metanephridial organs, and the original coelom body-cavity has degenerated to a pericardial cavity and gonads. Except for the pharynx, heart and gonads, the organs are enclosed in a membrane called an epicardium, which is surrounded by the jelly-like mesenchyme. Tunicates begin life in a mobile larval stages that resembles a a tadpole, later developing into a barrel-like, sedentary adult form.

Most tunicates are hermaphrodites. The eggs are kept inside their body until they hatch, while sperm is released into the water where it fertilizes other individuals when brought in with incoming water.

Some larval forms appear very much like primitive chordates or hemichordates with a notochord (primitive spinal cord). Superficially the larva resemble small tadpoles. Some forms have a calcereous spicule that may be preserved as a fossil. They have appeared from the Jurassic to the present, with one proposed Neoproterozoic form,

The larval stage ends when the tunicate finds a suitable rock to affix to and cements itself in place. The larval form is not capable of feeding, and is only a dispersal mechanism. Many physical changes occur to the tunicate's body, one of the most interesting being the digestion of the cerebral ganglion previously used to control movement. From this comes the common saying that the sea squirt "eats its own brain". In some classes, the adults remain pelagic (swimming or drifting in the open sea), although their larvae undergo similar metamorphoses to a higher or lower degree.

Sea anemone

Sea anemones are a group of water dwelling, predatory animals of the order Actiniaria; they are named after the anemone, a terrestrial flower. As cnidarians, sea anemones are closely related to corals, jellyfish, tube-dwelling anemones and Hydra.

A sea anemone is a small sac, attached to the bottom by an adhesive foot, called a basal disc, with a column shaped body ending in an oral disc. The mouth is in the middle of the oral disc, surrounded by tentacles armed with many cnidocytes, which are cells that function as a defense and as a means to capture prey. Cnidocytes contain cnidae, capsule-like organelles capable of everting, giving phylum Cnidaria its name . The cnidae that sting are called nematocysts. Each nematocyst contains a small vesicle filled with toxins—actinoporins—an inner filament and an external sensory hair. When the hair is touched, it mechanically triggers the cell explosion, a harpoon-like structure which attaches to organisms that trigger it, and injects a dose of poison in the flesh of the aggressor or prey. This gives the anemone its characteristic sticky feeling.

The poison is a mix of toxins, including neurotoxins, which paralyze the prey, which is then moved by the tentacles to the mouth/anus for digestion inside the gastrovascular cavity. Actinoporins have been reported as highly toxic to fish and crustaceans, which may be the natural prey of sea anemones. In addition to their role in predation, it has been suggested that actinoporins could act, when released in water, as repellents against potential predators. Certain clownfish are not affected by their host anemone's sting.

The internal anatomy of anemones is simple. There is a gastrovascular cavity (which functions as a stomach) with a single opening to the outside which functions as both a mouth and an anus: waste and undigested matter is excreted through the mouth/anus, which can be described as an incomplete gut. A primitive nervous system, without centralization, coordinates the processes involved in maintaining homeostasis as well as biochemical and physical responses to various stimuli. Anemones range in size from less than 1¼ cm (½ in) to nearly 2 m (6 ft) in diameter.They can have a range of ten tentacles to hundreds.

The muscles and nerves in anemones are much simpler than those of other animals. Cells in the outer layer (epidermis) and the inner layer (gastrodermis) have microfilaments grouped together into contractile fibers. These are not true muscles because they are not freely suspended in the body cavity as they are in more developed animals. Since the anemone lacks a skeleton, the contractile cells pull against the gastrovascular cavity, which acts as a hydrostatic skeleton. The stability for this hydrostatic skeleton is caused by the anemone shutting its mouth, which keeps the gastrovascular cavity at a constant volume, making it more rigid.

Mollusca

The molluscs (British spelling) or mollusks (American spelling) are members of the very large and diverse phylum of invertebrate animals known as Mollusca. There are some 112,000 species within this phylum. The scientific study of molluscs is known as malacology.

The word mollusc comes from the French mollusque, which originated from the Latin molluscus, meaning thin-shelled, from mollis, soft.

Molluscs range from minute snails and clams (micromollusks) to large organisms such as squid, cuttlefish and octopus, which are among the most neurologically-advanced invertebrates.

There are a wide variety of molluscs which are valued by humans as seafood or for their decorative shells. The edible species include many kinds of clams, snails, squid and octopuses.

The vast majority of molluscs live in marine environments, and many of them are found intertidally, in the shallow subtidal and on the continental shelf. Species of octopus and squid live throughout the ocean depths and some species of clam and limpet live in the abyssal depths of the oceans around hot vents.

Not all molluscs are marine: two taxonomic groups or classes, the bivalves and the gastropods, also contain freshwater species. Only the gastropods have representatives that live on land: the land snails and slugs.

Sponge

The sponges or poriferans (from Latin porus "pore" and ferre "to bear") are animals of the phylum Porifera,porifera translates to "Pore-bearer". They are primitive, sessile, mostly marine, water dwelling filter feeders that pump water through their bodies to filter out particles of food matter. Sponges represent the simplest of animals. With no true tissues (parazoa), they lack muscles, nerves, and internal organs. Their similarity to colonial choanoflagellates shows the probable evolutionary jump from unicellular to multicellular organisms. However, recent genomic studies suggest they are not the most ancient lineage of animals, but may instead by secondarily simplified.

There are over 5,000 modern species of sponges known, and they can be found attached to surfaces anywhere from the intertidal zone to as deep as 8,500 m (29,000 feet) or further. Though the fossil record of sponges dates back to the Neoproterozoic Era, new species are still commonly discovered.

Echinoderm

Echinoderms (Phylum Echinodermata, from the Greek for spiny skin) are a phylum of marine animals found at all ocean depths. Aside from the problematic Arkarua, the first definitive members of the phylum appeared near the start of the Cambrian period, and contains about 7,000 living species, making it the second largest grouping of deuterostomes, after the chordates; they are the largest phylum without freshwater or terrestrial representatives.

The Echinoderms are important both biologically and geologically: biologically because few other groupings are so abundant in the biotic desert of the deep sea, as well as the shallower oceans, and geologically as their ossified skeletons are major contributors to many limestone formations, and can provide valuable clues as to the geological environment. Further, it is held by some that the radiation of echinoderms was responsible for the Mesozoic revolution of marine life.

Two main subdivisions of Echinoderms are traditionally recognised: the more familiar, motile Eleutherozoa, which encompasses the Asteroidea (starfish), Ophiuroidea (brittle stars), Echinoidea (sea urchin and sand dollar) and Holothuroidea (sea cucumbers); and the sessile Pelmatazoa, which consist of the crinoids. Some crinoids, the feather stars, have secondarily re-evolved a free-living lifestyle.

A fifth class of Eleutherozoa consisting of just two species, the Concentricycloidea (sea daisies), were recently merged into the Asteroidea. The fossil record contains a host of other classes which do not appear to fall into any extant crown group.

Dinoflagellate

The dinoflagellates are a large group of flagellate protists. Most are marine plankton, but they are common in fresh water habitats as well. Their populations are distributed depending on temperature, salinity, or depth. About half of all dinoflagellates are photosynthetic, and these make up the largest group of eukaryotic algae aside from the diatoms. Being primary producers make them an important part of the aquatic food chain. Some species, called zooxanthellae, are endosymbionts of marine animals and protozoa, and play an important part in the biology of coral reefs. Other dinoflagellates are colorless predators on other protozoa, and a few forms are parasitic.

Dinoflagellates have a complex cell covering called an amphiesma, composed of flattened vesicles, called alveoli. In some forms, these support overlapping cellulose plates that make up a sort of armor called the theca. These come in various shapes and arrangements, depending on the species and sometimes stage of the dinoflagellate. Fibrous extrusomes are also found in many forms. Together with various other structural and genetic details, this organization indicates a close relationship between the dinoflagellates, Apicomplexa, and ciliates, collectively referred to as the alveolates.

The chloroplasts in most photosynthetic dinoflagellates are bound by three membranes, suggesting they were probably derived from some ingested algae, and contain chlorophylls a and c and either peridinin or fucoxanthin, as well as various other accessory pigments. However, a few have chloroplasts with different pigmentation and structure, some of which retain a nucleus. This suggests that chloroplasts were incorporated by several endosymbiotic events involving already colored or secondarily colorless forms. The discovery of plastids in Apicomplexa have led some to suggest they were inherited from an ancestor common to the two groups, but none of the more basal lines have them.

All the same, the dinoflagellate still consists of the more common organelles such as rough and smooth endoplasmic reticulum, Golgi apparatus, mitochondria, lipid and starch grains, and food vacuoles. Some have even been found with light sensitive organelle such as the eyespot or a larger nucleus containing a prominent nucleolus.

Diatom

Diatoms (Greek: διά (dia) = "through" + τέμνειν (temnein) = "to cut", i.e., "cut in half") are a major group of eukaryotic algae, and are one of the most common types of phytoplankton. Most diatoms are unicellular, although some form chains or simple colonies.

A characteristic feature of diatom cells is that they are encased within a unique cell wall made of silica (hydrated silicon dioxide) called a frustule. These frustules show a wide diversity in form, some quite beautiful and ornate, but usually consist of two asymmetrical sides with a split between them, hence the group name. Fossil evidence suggests that they originated during, or before, the early Jurassic Period. Diatom communities are a popular tool for monitoring environmental conditions, past and present, and are commonly used in studies of water quality.

Diatoms belong to a large group called the heterokonts, including both autotrophs ( golden algae, kelp) and heterotrophs (water moulds). Their yellowish-brown chloroplasts are typical of heterokonts, with four membranes and containing pigments such as fucoxanthin. Individuals usually lack flagella, but they are present in gametes and have the usual heterokont structure, except they lack the hairs (mastigonemes) characteristic in other groups. Most diatoms are non-motile, although some move via flagellation. As their relatively dense cell walls cause them to readily sink, planktonic forms in open water usually rely on turbulent mixing of the upper layers by the wind to keep them suspended in sunlit surface waters. Some species actively regulate their buoyancy with intracellular lipids to counter sinking.

Diatoms cells are contained within a unique silicate (silicic acid) cell wall comprised of two separate valves (or shells). The biogenic silica that the cell wall is composed of is synthesised intracellularly by the polymerisation of silicic acid monomers. This material is then extruded to the cell exterior and added to the wall. Diatom cell walls are also called frustules or tests, and their two valves typically overlap one other like the two halves of a petri dish. In most species, when a diatom divides to produce two daughter cells, each cell keeps one of the two valves and grows a smaller valve within it. As a result, after each division cycle the average size of diatom cells in the population gets smaller. Once such cells reach a certain minimum size, rather than simply divide vegetatively, they reverse this decline by forming an auxospore. This expands in size to give rise to a much larger cell, which then returns to size-diminishing divisions. Auxospore production is almost always linked to meiosis and sexual reproduction.

Tide pool

Tide pools are rocky pools by oceans that are filled with seawater. Tide pools can either be elephants,small and shallow or turtles and deep. The small ones are usually found far back on the shore and the large ones are found nearer to the ocean. Tide pools are formed as a high tide comes in over a rocky shore. Water fills depressions in the ground, which turn into isolated pools as the tide retreats. This process, repeated twice a day, replenishes the seawater in what otherwise might be a stagnant pool.

The area that is covered by high tide and exposed by low tide is called the intertidal zone, or foreshore. This area is often further divided into different zones based on the life forms that live there.

Seagrass

Seagrasses (or sea-grasses in British English) are flowering plants from one of four plant families (Posidoniaceae, Zosteraceae, Hydrocharitaceae, or Cymodoceaceae) which grow in marine, fully saline environments.

These unusual marine flowering plants are called seagrasses because the leaves are long and narrow and are very often green, and because the plants often grow in large "meadows" which look like grassland: in other words many of the species of seagrasses superficially resemble terrestrial grasses of the family Poaceae.

Because these plants must photosynthesize, they are limited to growing submerged in the photic zone, and most occur in shallow and sheltered coastal waters anchored in sand or mud bottoms. They undergo pollination while submerged and complete their entire life cycle underwater. There are about sixty species worldwide.

Seagrasses form extensive beds or meadows, which can be either monospecific (made up of one species) or multispecific (where more than one species co-exist). In temperate areas, usually one or a few species dominate (like the eelgrass Zostera marina in the North Atlantic), whereas tropical beds usually are more diverse, with up to thirteen species recorded in the Philippines.

Seagrass beds are highly diverse and productive ecosystems, and can harbor hundreds of associated species from all phyla, for example juvenile and adult fish, epiphytic and free-living macroalgae and microalgae, mollusks, bristle worms, and nematodes. Few species were originally considered to feed directly on seagrass leaves (partly because of their low nutritional content), but scientific reviews and improved working methods have shown that seagrass herbivory is a highly important link in the food chain, with hundreds of species feeding on seagrasses worldwide, including dugongs, manatees, fish, geese, swans, sea urchins and crabs.

Seagrasses are sometimes labeled ecosystem engineers, because they partly create their own habitat: the leaves slow down water-currents increasing sedimentation, and the seagrass roots and rhizomes stabilize the seabed. Their importance for associated species is mainly due to provision of shelter (through their three-dimensional structure in the water column), and for their extraordinarily high rate of primary production. As a result, seagrasses provide coastal zones with a number of ecosystem goods and ecosystem services, for instance fishing grounds, wave protection, oxygen production and protection against coastal erosion.

Seaweed

Seaweeds are any of a large number of marine plants in the category of benthic algae. They are macroscopic and multicellular, in contrast with most other algae. Seaweeds are often found in the seashore biome.

Seaweeds consist of several groups of multicellular algae: the red algae, green algae, and brown algae. As these three groups are not thought to have a common multicellular ancestor, the seaweeds are a paraphyletic group. In addition, tuft-forming bluegreen algae (Cyanobacteria) are sometimes considered as seaweeds.

Seaweeds are popularly described as plants, but only red and green algae belong to the kingdom Plantae. They should not be confused with aquatic plants such as seagrasses (which are vascular plants).

Aquatic plant

Aquatic plants — also called hydrophytic plants or hydrophytes — are plants that have adapted to living in or on aquatic environments. Because living on or under water surface requires numerous special adaptations, aquatic plants can only grow in water or permanently saturated soil. Aquatic vascular plants can be ferns or angiosperms (from both monocot and dicot families).

Seaweeds are not vascular plants but multicellular marine algae, and therefore not typically included in the category of aquatic plants. As opposed to plants types such as mesophytes and xerophytes, hydrophytes do not have a problem in retaining water due to the abundance of water in its environment. This means the plant has less need to regulate transpiration (indeed, the regulation of transpiration would require more energy than the possible benefits incurred.)

Brown algae

The Phaeophyceae or brown algae, (singular: alga) is a large group of mostly marine multicellular algae, including many seaweeds of colder Northern Hemisphere waters. They play an important role in marine environments both as food, and for the habitats they form. For instance Macrocystis, a member of the Laminariales or kelps, may reach 60 m in length, and forms prominent underwater forests. Another example is Sargassum, which creates unique habitats in the tropical waters of the Sargasso Sea. This is one of the few areas where a large biomass of brown algae may be found in tropical waters. Many brown algae such as members of the order Fucales are commonly found along rocky seashores. Some members of the division are used as food for humans.

Brown algae belong to a very large group, the Heterokontophyta, a eukaryotic group of organisms distinguished most prominently by having chloroplasts surrounded by four membranes, suggesting an origin from a symbiotic relationship between a basal eukaryote and another eukaryotic organism. Most brown algae contain the pigment fucoxanthin, which is responsible for the distinctive greenish-brown color that gives them their name. Brown algae are unique among heterokonts in developing into multicellular forms with differentiated tissues, but they reproduce by means of flagellate spores, which closely resemble other heterokont cells. Genetic studies show their closest relatives to be the yellow-green algae.

Phaeophyta first appear in the fossil record in the Mesozoic, possibly as early as the Jurassic. Their occurrence as fossils is rare due to their generally soft-bodied habit, and scientists continue to debate the identification of some finds. Other algae groups, such as the red algae and green algae have a number of calcareous members, which are more likely to leave evidence in the fossil record than the soft bodies of the brown algae. Miocene fossils of a soft-bodied brown macro algae, Julescrania, have been found well-preserved in Monterey Formation diatomites, but few other certain fossils, particularly of older specimens are known in the fossil record.

Ocean

An ocean (from Ωκεανός, Okeanos (Oceanus) in Greek) is a major body of saline water, and a principal component of the hydrosphere. Approximately 71% of the Earth's surface (an area of some 361 million square kilometers) is covered by ocean, a continuous body of water that is customarily divided into several principal oceans and smaller seas. More than half of this area is over 3,000 meters (9,800 ft) deep. Average oceanic salinity is around 35 parts per thousand (ppt) (3.5%), and nearly all seawater has a salinity in the range of 31 to 38 ppt.

Though generally recognized as several 'separate' oceans, these waters comprise one global, interconnected body of salt water often referred to as the World Ocean or global ocean.This concept of a global ocean as a continuous body of water with relatively free interchange among its parts is of fundamental importance to oceanography.The major oceanic divisions are defined in part by the continents, various archipelagos, and other criteria: these divisions are (in descending order of size) the Pacific Ocean, the Atlantic Ocean, the Indian Ocean, the Southern Ocean (which is sometimes subsumed as the southern portions of the Pacific, Atlantic, and Indian Oceans), and the Arctic Ocean (which is sometimes considered a sea of the Atlantic). The Pacific and Atlantic may be further subdivided by the equator into northerly and southerly portions. Smaller regions of the oceans are called seas, gulfs, bays and other names. There are also some smaller bodies of saltwater that are on land and not interconnected with the World Ocean, such as the Aral Sea, and the Great Salt Lake – though they may be referred to as 'seas', they are actually salt lakes.

Geologically, an ocean is an area of oceanic crust covered by water. Oceanic crust is the thin layer of solidified volcanic basalt that covers the Earth's mantle where there are no continents. From this perspective, there are three oceans today: the World Ocean and the Caspian and Black Seas, the latter two having been formed by the collision of Cimmeria with Laurasia. The Mediterranean Sea is very nearly a discrete ocean, being connected to the World Ocean through the Strait of Gibraltar, and indeed several times over the last few million years movement of the African continent has closed the strait off entirely. The Black Sea is connected to the Mediterranean through the Bosporus, but this is in effect a natural canal cut through continental rock some 7,000 years ago, rather than a piece of oceanic sea floor like the Strait of Gibraltar.

Coralline algae

Coralline algae are red algae in the Family Corallinaceae of the order Corallinales characterized by a thallus that is hard as a result of calcareous deposits contained within the cell walls. Unattached specimens (maerl, rhodoliths) may form relatively smooth compact balls to warty or fruticose thalli. Many are typically encrusting and rock-like, found in tropical marine waters all over the world. They play an important role in the ecology of coral reefs. Colors are most typically pink or some other shade of red, but may be purple, yellow, blue, white or gray-green. Sea urchins, parrot fish, limpets (molluscs) and chitons molluscs feed on coralline algae.

A close look at almost any intertidal rocky shore or coral reef will reveal an abundance of pink to pinkish-grey patches, splashed as though by a mad painter over rock surfaces. These patches of pink paint are actually living algae: crustose coralline red algae. The red algae belong to the division Rhodophyta, within which the coralline algae form a distinct, exclusively marine order, the Corallinales. Coralline algae are widespread in all of the world's oceans, where they often cover close to 100% of rocky substrata. Many are epiphytic (grow on other algae or marine angiosperms), or epizoic (grow on animals), and some are even parasitic on other corallines. Despite their ubiquity, the coralline algae are poorly known by ecologists, and even by specialist phycologists (people who study algae). For example, a recent book on the seaweeds of Hawai'i does not include any crustose coralline algae even though corallines are quite well studied there and dominate many marine areas.

Red algae

The red algae (Rhodophyta, IPA, from Greek: (rhodon) rose + (phyton) = plant, thus red plant) are a large group, about 5,000–6,000 species of mostly multicellular, marine algae, including many notable seaweeds. Other references indicate 10,000 species. Most of the coralline algae, which secrete calcium carbonate and play a major role in building coral reefs, belong here. Red algae such as dulse (Palmaria palmata) and laver (nori/gim) are a traditional part of European and Asian cuisine and are used to make other products like agar, carrageenans and other food additives.

Algae

Algae (sing. alga) are a large and diverse group of simple plant-like organisms, ranging from unicellular to multicellular forms. The largest and most complex marine forms are called seaweeds. They are considered "plant-like" because of their photosynthetic ability, and "simple" because they lack the distinct organs of higher plants such as leaves and vascular tissue. Though the prokaryotic Cyanobacteria (commonly referred to as Blue-green algae) were traditionally included as "algae" in older textbooks, many modern sources regard this as outdated and restrict the term algae to eukaryotic organisms.All true algae therefore have a nucleus enclosed within a membrane and chloroplasts bound in one or more membranes.Algae constitute a paraphyletic and polyphyletic group: they do not represent a single evolutionary direction or line, but a level or grade of organization that may have developed several times in the early history of life on Earth.

Algae lack leaves, roots, and other organs that characterize higher plants. They are distinguished from protozoa in that they are photosynthetic. Many are photoautotrophic, although some groups contain members that are mixotrophic, deriving energy both from photosynthesis and uptake of organic carbon either by osmotrophy, myzotrophy, or phagotrophy. Some unicellular species rely entirely on external energy sources and have reduced or lost their photosynthetic apparatus.

All algae have photosynthetic machinery ultimately derived from the cyanobacteria, and so produce oxygen as a byproduct of photosynthesis, unlike other photosynthetic bacteria such as purple and green sulfur bacteria.

Golden algae

The golden algae or chrysophytes are a large group of heterokont algae, found mostly in freshwater. Originally they were taken to include all such forms except the diatoms and multicellular brown algae, but since then they have been divided into several different groups based on pigmentation and cell structure. They are now usually restricted to a core group of closely related forms, distinguished primarily by the structure of the flagella in motile cells, also treated as an order Chromulinales. It is possible membership will be revised further as more species are studied in detail. They come in a variety of morphological types, originally treated as separate orders or families.

Most members are unicellular flagellates, with either two visible flagella, as in Ochromonas, or sometimes one, as in Chromulina. The Chromulinales as first defined by Pascher in 1910 included only the latter type, with the former treated as the order Ochromonadales. However, structural studies have revealed that short second flagellum or at least a second basal body is always present, so this is no longer considered a valid distinction. Most of these have no cell covering. Some have loricae or shells, such as Dinobryon, which is sessile and grows in branched colonies. Most forms with silicaceous scales are now considered a separate group, the synurids, but a few belong among the Chromulinales proper, such as Paraphysomonas.

Some members are generally amoeboid, with long branching cell extensions, though they pass through flagellate stages as well. Chrysamoeba and Rhizochrysis are typical of these. There is also one species, Myxochrysis paradoxa, which has a complex life cycle involving a multinucleate plasmodial stage, similar to those found in slime moulds. These were originally treated as the order Chrysamoebales. The superficially similar Rhizochromulina was once included here, but is now given its own order based on differences in the structure of the flagellate stage.

Other members are non-motile. Cells may be naked and embedded in mucilage, such as Chrysosaccus, or coccoid and surrounded by a cell wall, as in Chrysosphaera. A few are filamentous or even parenchymatous in organization, such as Phaeoplaca. These were included in various older orders, most of the members of which are now included in separate groups. Hydrurus and its allies, freshwater genera which form branched gelatinous filaments, are often placed in the separate order Hydrurales but may belong here.

Chrysophytes were once considered to be a specialized form of cyanobacteria containing the golden-yellow pigment, fucoxanthin. Because many of these organisms had a silica capsule, they have a relatively complete fossil record, allowing modern biologists to confirm that they are, in fact, not derived from cyanobacteria, but rather an ancestor that did not possess the capability to photosynthesize. Many of the chrysophyta precursor fossils entirely lacked any type of photosynthesis-capable pigment. Most biologists believe that the chrysophytes obtained their ability to photosynthesis from an endosymbiotic relationship with fucoxanthin-containing cyanobacteria.

Stromatolite

Stromatolites (from Greek strōma, mattress, bed, stratum, and lithos, rock) are layered accretionary structures formed in shallow water by the trapping, binding, and cementation of sedimentary grains by biofilms of microorganisms, especially cyanobacteria (commonly known as blue-green algae). A variety of stromatolite morphologies exist including conical, stratiform, branching, domal, and columnar types. Stromatolites occur widely in the fossil record of the Precambrian, but are rare today. Very few ancient stromatolites contain fossilized microbes. While features of some stromatolites are suggestive of biological activity, others possess features that are more consistent with "abiotic" (non-organic) precipitation. Finding reliable ways to distinguish between biologically-formed and abiotic (non-biological) "stromatolites" is an active area of research in geology.

Stromatolites were much more abundant on the planet in Precambrian times. While older, Archean fossil remains are presumed to be colonies of single-celled blue-green bacteria, younger (that is, Proterozoic) fossils may be primordial forms of the eukaryote chlorophytes (that is, green algae). One genus of stromatolite very common in the geologic record is Collenia. The earliest stromatolite of confirmed microbial origin dates to 2,724 million years ago.

Prior to 2.4 billion years ago, the earth's atmosphere was rich in carbon dioxide. However, the Precambrian air lacked the oxygen that sustains the complex multicellular life that has evolved since the "Cambrian explosion". 540Ma. Stromatolites in the fossil record decline sharply in both diversity and number during the late Proterozoic eon, although they are present, but not common, in Paleozoic era strata. Today, stromatolites are quite uncommon in marine environments, and thus are called "living fossils."

Their former abundance may be because there were no burrowing or grazing animals back during the Precambrian to destabilize sediments and consume growing microbial mats, thereby favoring the preservation of these microbialites. Also, changing chemical conditions in the ocean during this time could be responsible for the precipitation of non-biological stromatolites through the growth of tiny crystals.

While prokaryotic cyanobacteria themselves reproduce asexually through cell division, they were instrumental in priming the environment for the evolutionary development of more complex eukaryotic organisms. Cyanobacteria are thought to be largely responsible for increasing the amount of oxygen in the primeval earth's atmosphere through their continuing photosynthesis.

Cyanobacteria use water, carbon dioxide, and sunlight to create their food. The byproducts of this process are oxygen and calcium carbonate (lime). A layer of mucus often forms over mats of cyanobacterial cells. In modern microbial mats, debris from the surrounding habitat can become trapped within the mucus, which can be cemented together by the calcium carbonate to grow thin laminations of limestone. These laminations can accrete over time, resulting in the banded pattern common to stromatolites. The domal morphology of biological stromatolites is the result of the vertical growth necessary for the continued infiltration of sunlight to the organisms for photosynthesis.

Modern stromatolites are mostly found in hypersaline lakes and marine lagoons where extreme conditions due to high saline levels exclude animal grazing.

Cyanobacteria

Cyanobacteria, also known as blue-green algae, blue-green bacteria or Cyanophyta, is a phylum of bacteria that obtain their energy through photosynthesis. The name "cyanobacteria" comes from the color of the bacteria (Greek: κυανός (kyanós) = blue). They are a significant component of the marine nitrogen cycle and an important primary producer in many areas of the ocean, but are also found on land.

Stromatolites of fossilized oxygen-producing cyanobacteria have been found from 2.8 billion years ago.The ability of cyanobacteria to perform oxygenic photosynthesis is thought to have converted the early reducing atmosphere into an oxidizing one, which dramatically changed the life forms on Earth and provoked an explosion of biodiversity. Chloroplasts in plants and eukaryotic algae have evolved from cyanobacteria.

Marine invertebrates

The term "marine invertebrates" is used to describe animals found in a marine environment which are invertebrates: lacking a notochord. In order to protect themselves, they may have evolved a shell or a hard exoskeleton, but this is not always the case.

As on land and in the air, invertebrates make up a huge portion of all life in the sea. Invertebrate sea life includes:

• Bryozoa, also known as moss animals or sea mats;


• Cnidaria, such as jellyfish, sea anemones and corals;


• Crustaceans, such a such as lobsters, crabs, shrimp, crayfish and barnacles;


• Ctenophora: sea worms including flatworms, ribbon worms, annelids, Sipuncula, Echiura, Chaetognatha, and the phoronids;


• Echinoderms, including starfish, brittle stars, sea urchins, sand dollars, sea cucumbers, and crinoids;


• Mollusca, including shellfish, squid, octopus;


• Sponges;


• Tunicates, also known as sea squirts.

Zooplankton

Zooplankton are the heterotrophic (sometimes detritivorous) component of the plankton that drift in the water column of oceans, seas, and bodies of fresh water. The name is derived from the Greek terms, ζῴον ("zoon") meaning "animal", and πλαγκτος ("planktos") meaning "wanderer" or "drifter". Many zooplankton are too small to be individually seen with the unaided eye.

Zooplankton is a broad categorisation spanning a range of organism sizes that includes both small protozoans and large metazoans. It includes holoplanktonic organisms whose complete life cycle lies within the plankton, and meroplanktonic organisms that spend part of their life cycle in the plankton before graduating to either the nekton or a sessile, benthic existence.

Ecologically important protozoan zooplankton groups include the foraminiferans, radiolarians and dinoflagellates (the latter are often mixotrophic). Important metazoan zooplankton include cnidarians such as jellyfish and the Portuguese Man o' War; crustaceans such as copepods and krill; chaetognaths (arrow worms); molluscs such as pteropods; and chordates such as salps and juvenile fish. This wide phylogenetic range includes a similarly wide range in feeding behaviour: filter feeding, predation and symbiosis with autotrophic phytoplankton as seen in corals. Zooplankton feed on bacterioplankton, phytoplankton, other zooplankton (sometimes cannibalistically), detritus (or marine snow) and even nektonic organisms.

Through their consumption and processing of phytoplankton (and other food sources), zooplankton play an important role in aquatic food webs, both as a resource for consumers on higher trophic levels and as a conduit for packaging the organic material in the biological pump.

Fish

Fish are aquatic vertebrates that are typically ectothermic (previously cold-blooded), covered with scales, and equipped with two sets of paired fins and several unpaired fins. Fish are abundant in the sea and in fresh water, with species being known from mountain streams char and gudgeon as well as in the deepest depths of the ocean gulpers and anglerfish.

They are of tremendous importance as food for people around the world, either collected from the wild. fishing or farmed in much the same way as cattle or chickens . Fish are also exploited for recreation, through angling and fishkeeping, and are commonly exhibited in public aquaria. Fish have an important role in many cultures through the ages, ranging as widely as deities and religious symbols to subjects of books and popular movies.

Marine reptile

Marine reptiles are reptiles which have become secondarily adapted for an aquatic or semi-aquatic life in a marine environment.

The earliest marine reptiles arose in the Permian period during the Paleozoic era. During the Mesozoic era, many groups of reptiles became adapted to life in the seas, including such familiar clades as the ichthyosaurs, plesiosaurs, placodonts, and mosasaurs.

After the first mass extinction at the end of the Cretaceous period, marine reptiles are less numerous, consisting largely of sea turtles, sea snakes, Marine Iguanas and some species of crocodylians.

Some marine reptiles, such as ichthyosaurs and mosasaurs, rarely if ever ventured onto land and gave birth in the water. Others, such as sea turtles and saltwater crocodiles, still return to shore to lay their eggs.

Adaptations

Since mammals originally evolved on land, their spines are optimized for running, allowing for up-and-down but only little sideways motion. Therefore, marine mammals typically swim by moving their spine up and down. By contrast, fish normally swim by moving their spine sideways. For this reason, fish mostly have vertical caudal (tail) fins, while marine mammals have horizontal caudal fins.

Some of the primary differences between marine mammals and other marine life are:

• Marine mammals breathe air, while most other marine animals extract oxygen from water.


• Marine mammals have hair. Cetaceans have little or no hair, usually a very few bristles retained around the head or mouth. All members of the Carnivora have a coat of fur or hair, but it is far thicker and more important for thermoregulation in sea otters and polar bears than in seals or sea lions. Thick layers of fur contribute to drag while swimming, and slow down a swimming mammal, giving it a disadvantage in speed.


• Marine mammals have thick layers of blubber used to insulate their bodies and prevent heat loss. Sea otters and polar bears are exceptions, relying more on fur and behavior to stave off hypothermia.


• Marine mammals give birth. Most marine mammals give birth to one calf or pup at a time.


• Marine mammals feed off milk as young. Maternal care is extremely important to the survival of offspring that need to develop a thick insulating layer of blubber. The milk from the mammary glands of marine mammals often exceeds 40-50% fat content to support the development of blubber in the young.


• Marine mammals maintain a high internal body temperature. Unlike most other marine life, marine mammals carefully maintain a core temperature much higher than their environment. Blubber, thick coats of fur, bubbles of air between skin and water, countercurrent exchange, and behaviors such as hauling out, are all adaptations that aid marine mammals in retention of body heat.

The polar bear spends a large portion of its time in a marine environment, albeit a frozen one. When it does swim in the open sea it is extremely proficient and has been shown to cover 74 km in a day. For these reasons, some scientists regard it as a marine mammal.

Marine mammal

Marine mammals are a diverse group of roughly 120 species of mammal that are primarily ocean-dwelling or depend on the ocean for food. They include the cetaceans (whales, dolphins, and porpoises), the sirenians (manatees and dugong), the pinnipeds (true seals, eared seals and walrus), and several otters (the sea otter and marine otter). The polar bear is also usually grouped with the marine mammals.

Marine mammals evolved from land dwelling ancestors and share several adaptive features for life at sea such as generally large size, hydrodynamic body shapes, modified appendages and various thermoregulatory adaptations. Different species are, however, adapted to marine life to varying degrees. The most fully adapted are the cetaceans and the sirenians, whose entire life cycle takes place under water, whereas the other groups spend at least some time on land.

Despite the fact that marine mammals are highly recognizable charismatic megafauna, many populations are vulnerable or endangered due to a history of commercial exploitation for blubber, meat, ivory and fur. Most species are currently protected from commercial exploitation.

Oceanic Habitats

Reefs

comprise some of the densest and most diverse habitats in the world. The best-known types of reefs are tropical coral reefs which exist in most tropical waters; however, reefs can also exist in cold water. Reefs are built up by corals and other calcium-depositing animals, usually on top of a rocky outcrop on the ocean floor. Reefs can also grow on other surfaces, which has made it possible to create artificial reefs. Coral reefs also support a huge community of life, including the corals themselves, their symbiotic zooxanthellae, tropical fish and many other organisms.

Much attention in marine biology is focused on coral reefs and the El Niño weather phenomenon. In 1998, coral reefs experienced a "once in a thousand years" bleaching event, in which vast expanses of reefs across the Earth died because sea surface temperatures rose well above normal. Some reefs are recovering, but scientists say that 58% of the world's coral reefs are now endangered and predict that global warming could exacerbate this trend.

Oceanic trench

The oceanic trenches are hemispheric-scale long but narrow topographic depressions of the sea floor. They are also the deepest parts of the ocean floor.

Trenches define one of the most important natural boundaries on the Earth’s solid surface, that between two lithospheric plates. There are three types of lithospheric plate boundaries: divergent (where lithosphere and oceanic crust is created at mid-ocean ridges), convergent (where one lithospheric plate sinks beneath another and returns to the mantle), and transform (where two lithospheric plates slide past each other). Trenches are the spectacular and distinctive morphological features of convergent plate boundaries. Plates move together along convergent plate boundaries at convergence rates that vary from a few millimeters to ten or more centimeters per year.

A trench marks the position at which the flexed, subducting slab begins to descend beneath another lithospheric slab. Trenches are generally parallel to a volcanic island arc, and trenches about 200 km from a volcanic arc. Oceanic trenches typically extend 3 to 4 km (1.9 to 2.5 mi) below the level of the surrounding oceanic floor. The deepest ocean depth to be sounded is in the Challenger Deep of the Mariana Trench at a depth of 10,911 m (35,798 ft) below sea level. Oceanic lithosphere disappears into trenches at a global rate of about a tenth of a square meter per second.

Filled trenches

The composition of the inner trench slope and a first-order control on trench morphology is determined by sediment supply. Active accretionary prisms are common for trenches near continents where large rivers or glaciers reach the sea and supply great volumes of sediment which naturally flow to the trench. These filled trenches are confusing because in a plate tectonic sense they are indistinguishable from other convergent margins but lack the bathymetric expression of a trench.

The Cascadia margin of the northwest USA is a filled trench, the result of sediments delivered by the rivers of the NW USA and SW Canada. The Lesser Antilles convergent margin shows the importance of proximity to sediment sources for trench morphology. In the south, near the mouth of the Orinoco River, there is no morphological trench and the forearc plus accretionary prism is almost 500 km wide. The accretionary prism is so large that it forms the islands of Barbados and Trinidad. Northward the forearc narrows, the accretionary prism disappears, and only north of 17°N the morphology of a trench is seen. In the extreme north, far away from sediment sources, the Puerto Rico Trench is over 8600 m deep and there is no active accretionary prism. A similar relationship between proximity to rivers, forearc width, and trench morphology can be observed from east to west along the Alaskan-Aleutian convergent margin. The convergent plate boundary offshore Alaska changes along its strike from a filled trench with broad forearc in the east (near the coastal rivers of Alaska) to a deep trench with narrow forearc in the west (offshore the Aleutian islands). Another example is the Makran convergent margin offshore Pakistan and Iran, which is a trench filled by sediments from the Tigris-Euphrates and Indus rivers. Thick accumulations of turbidites along a trench can be supplied by down-axis transport of sediments that enter the trench 1000-2000 km away, as is found for the Peru-Chile Trench south of Valparaíso and for the Aleutian Trench. Convergence rate can also be important for controlling trench depth, especially for trenches near continents, because slow convergence causes the capacity of the convergent margin to dispose of sediment to be exceeded.

There an evolution in trench morphology can be expected as oceans close and continents converge. While the ocean is wide, the trench may be far away from continental sources of sediment and so may be deep. As the continents approach each other, the trench may become filled with continental sediments and become shallower. A simple way to approximate when the transition from subduction to collision has occurred is when the plate boundary previously marked by a trench is filled enough to rise above sea level.

Intertidal and shore

Intertidal zones, those areas close to shore, are constantly being exposed and covered by the ocean's tides. A huge array of life lives within this zone.

Shore habitats span from the upper intertidal zones to the area where land vegetation takes prominence. It can be underwater anywhere from daily to very infrequently. Many species here are scavengers, living off of sea life that is washed up on the shore. Many land animals also make much use of the shore and intertidal habitats. A subgroup of organisms in this habitat bores and grinds exposed rock through the process of bioerosion.

How oceanic factors affect distribution of various organisms

An active research topic in marine biology is to discover and map the life cycles of various species and where they spend their time. Marine biologists study how the ocean currents, tides and many other oceanic factors affect ocean lifeforms, including their growth, distribution and well-being. This has only recently become technically feasible with advances in GPS and newer underwater visual devices.

Most ocean life breeds in specific places, nests or not in others, spends time as juveniles in still others, and in maturity in yet others. Scientists know little about where many species spent different parts of their life cycles. For example, it is still largely unknown where sea turtles travel. Tracking devices do not work for some life forms, and the ocean is not friendly to technology.