Annelid

The annelids, collectively called Annelida (from Latin anellus "little ring"), are a large phylum of animals comprising the segmented worms, with about 15,000 modern species including the well-known earthworms and leeches. They are found in most wet environments, and include many terrestrial, freshwater, and especially marine species (such as the polychaetes), as well as some which are parasitic or mutualistic. They range in length from under a millimeter to over 3 meters (the seep tube worm Lamellibrachia luymesi).

Annelids are bilaterally symmetric and triploblastic protostomes with a coelom (which makes them coelomates), closed circulatory system and true segmentation. Their segmented bodies and coelom have given them evolutionary advantages over other worms. Oligochaetes and polychaetes typically have spacious coeloms; in leeches, the coelom is filled in with tissue and reduced to a system of narrow canals; archiannelids may lack the coelom entirely. The coelom is divided into a sequence of compartments by walls called septa.

In the most general forms each compartment corresponds to a triple segment of the body, which also includes a portion of the nervous and (closed) circulatory systems, allowing it to function relatively independently. The closed circulatory system consists of networks of vessels containing blood with oxygen-carrying hemoglobin. Dorsal and ventral vessels are connected by segmental pairs of vessels. The dorsal vessel and five pairs of vessels that circle the esophagus of an earthworm are muscular and pump blood through the circulatory system.

Tiny blood vessels are abundant in the earthworm's skin, which function as its respiratory organ. Each segment (metamere) is marked externally by one or more rings, called annuli. Each segment also has an outer layer of circular muscle underneath a thin cuticle and epidermis, and a system of longitudinal muscles. In earthworms and in daria the longitudinal muscles are strengthened by collagenous lamellae; the leeches have a double layer of muscles between the outer circulars and inner longitudinals. In most forms they also carry a varying number of bristles, called setae, and among the polychaetes a pair of appendages, called parapodia.

Anterior to the true segments lies the prostomium and peristomium, which carries the mouth, and posterior to them lies the pygidium, where the anus is located. The digestive tract is quite variable but is usually specialized. For example, in some groups (notably most earthworms) it has a typhlosole (to increase surface area) along much of its length. Different species of annelids have a wide variety of diets, including active and passive hunters, scavengers, filter feeders, direct deposit feeders which simply ingest the sediments, and blood-suckers.

The vascular system and the nervous system are separate from the digestive tract. The vascular system includes a dorsal vessel conveying the blood toward the front of the worm, and a ventral longitudinal vessel which conveys the blood in the opposite direction. The two systems are connected by a vascular sinus and by lateral vessels of various kinds, including in the true earthworms, capillaries on the body wall.

The nervous system has a nerve cord from which lateral nerves come in contact with each segment. Every segment has an autonomy; however, they unite to perform as a single body for functions such as locomotion. Growth in many groups occurs by replication of individual segmental units, in others the number of segments is fixed in early development.